When Billy Dilley's science fair project the Cheeserator gets struck by lightning and goes haywire, Billy, his lab partners Zeke and Marsha, and his pet rat Anaximander, find themselves stuck in a strange underground land called Subterranea-Tania.

On their second day in Subterranea-Tania, Billy, Zeke, and Marsha find their way back to the Cheeserator after getting kidnapped by a vicious Pterosaur. After getting back to the Cheeserator, the trio builds a house made out of extra parts for safety.

Zeke, the King of Calzones, must defend his title when a rival chef enters the picture.